JFW Science and Professional Activities

Below is an informal summary of JFW’s educational and professional history.   Although this posting is not related to photography, it is a convenient way for me to maintain an active venue for reporting this work.   This site replaces a link (no longer active) to JFW’s research activities on the University of Delaware College of Earth, Ocean, and Environment web site (updated August 2022).


Dept. of Geological Sciences, College of Earth, Ocean, and Environment, Univ. Delaware,

Professor, Sept. 1986-present

Research Professor, George Washington University, 2003-

CHAIR, Dept. of Geology, 1996-2001, including sabbatical (acting chair, 1992)

Associate Prof., 1978 – 1985.

Assistant Prof., 1972-1978.

Research Associate, Invert. Paleontology, Los Angeles Co. Museum of Natural History, 2/86 – present

NASA-AMES Research Center Summer Faculty Fellow, 1974, 1975

Sabbatical leaves 1978; 1985; 1993; 2000-2001; 2007-2008



CO-CONVENOR, 1982 Geol. Society of America Symposium on Amino acid geochemistry and aminostratigraphy, New Orleans.

1981-1984  Organizer of international Interlaboratory Comparison Project – all active amino acid geochemistry laboratories.    See Wehmiller, 1984 (Quaternary Research)

Invited Participant in 1985 United Nations-CCOP Workshop on Dating Young Sediments, Beijing.

CO-CONVENOR, 1987 Geol. Society of America Penrose Conference on Late Pleistocene Sea Levels, Bermuda.

Invited Participant, 1989 International Geological Congress, Workshop on Biomineralization, Washington, D. C.

Theme-session organizer, 1990 Geological Society of America Meeting, Dallas.  16-paper session on the applications of amino acid racemization in geochronology, stratigraphy, and geochemistry.

International Geological Correlation Project 274: Coastal Quaternary Evolution: 

Theme leader for geochronology (liaison between radiocarbon labs and developing country coastal geologists); Co-editor, US Working Group final report: Quaternary Coastal Systems of the U. S. (published by Society of Economic Paleontologists and Mineralogists, late 1992 (Fletcher and Wehmiller)

FELLOWSHIP,  1995-1996 University of Delaware Center for Advanced Study 

CONVENOR, 1997 Northeastern Section Geological Society of America Workshop on Aminostratigraphy, held at University of Delaware, March 1997

CO-CONVENOR, 1998 Perspectives on Amino Acid and Protein Geochemistry: Symposium to Honor P. E. Hare, Carnegie Inst. of Washington. American Geophysical Union, Washington, DC.  April 1998.  (Co-editor of Proceedings Volume)

CO-CONVENOR, 1999 Assateague Shore and Shelf Workshop, Lewes, DE April 1999; maintenance of ASSW web site 2002-

Member, Editorial Advisory Board, Quaternary Science Reviews, 1991-1996; Quaternary Geochronology 2005 to ~2010 (?)

SERVICE on many University Committees (1990-1991 CHAIR of University Promotion and Tenure Committee);  Chair of Geology  Dept. P&T Committee, 1986-1991; Department Safety Committee Chair ~1990-2010; Dept. Advisory Committee;  Univ. Faculty Senate, 2004-2006; University Sustainability Task Force, 2006-2010. Regular reviewer for numerous journals such as Quaternary Research, Geological Society of America, Nature, and Geochimica Cosmochimica Acta, and for U. S. National Science Foundation.

Environmental Science Program, Univ. of Del. 1992- (Geol. Dept. rep. involved in undergrad. and grad. programs in Environmental Science)

Program Director Undergraduate Studies, College of Marine Studies, January 2004-January 2005

2004-present: Member of Franklin Institute Committee on Science and the Arts, Earth and Environmental Science Cluster

2006 Department committees:  Safety (Chair); Promotion and Tenure; University Senate; Search committee (hydrogeology)

Cutting Edge Teaching Workshop, June 2008: Teaching with ice core data (Penn State AMQUA meeting)

College of Earth, Ocean, and Environment P&T committee Chair  2009-2010

UD Sustainability Task Force

Fellow, UD Center for Energy and Environmental Policy, 2009-2012

2010 Denver GSA NOAA Database workshop NSF funding managed by JFW

2011  Bern Switzerland INQUA workshop for AAR data management $$  NSF funding for US participants

My official retirement began in January 2013 but was prefaced with a one-year sabbatical after completing my last teaching class in December 2011.  Leading up to retirement, I spent a great deal of time planning for the distribution of research samples that we had collected during the previous 40 years.  This effort also included a collaboration with several colleagues to establish an archive for Amino Acid Racemization (AAR) data at the NOAA Paleoclimatology site (see links below).  Beginning in January 2013, I devoted large amounts of time to the organization of our research collection and the physical transportation or shipment of samples to several institutions around the country (Los Angeles County Museum of Natural History; Paleontological Research Institution, Ithaca, NY;  Florida Museum of Natural History, Gainesville, FL; The Academy of Natural Sciences of Drexel University, Philadelphia; and the Delaware and North Carolina Geological Surveys), an effort that required about three years to complete.   Several efforts to synthesize data and to explain our data management procedures also occurred during this time (Wehmiller et al., 2015; Wehmiller and Pellerito, 2015a, 2015b).  


A fortuitous an unexpected combination of events starting in 2014 led to a major project that has resulting in a very satisfying publication (Wehmiller et al., 2021). I credit my colleague Kelvin Ramsey of the Delaware Geological Survey with the insight to realize that AAR data would be useful in understanding the links between the offshore and onshore Quaternary stratigraphy of the mid-Atlantic coastal region, thereby inviting me to participate in this multi-year project. Only in retrospect do I realize that my participation in this project would have been difficult had I had all the “normal” responsibilities associated with a full-time faculty position. Ramsey was responsible for managing an offshore sand resource project funded by the Bureau of Ocean Energy Management (BOEM), motivated by the need to assess these resources post- Hurricane Sandy.   The on-going offshore geophysical work of Laura Brothers (USGS, Woods Hole) and her colleagues was a major component of this project, allowing us to understand the geochronologic relation of offshore and onshore units, and even gain insights into the late Quaternary history of local barrier islands.  The mid-Atlantic results also have a counterpart in a related BOEM project for the SE US (Wehmiller et al., 2019.)  These offshore studies will hopefully contribute insights not only related to sand resources but also for the understanding of late Quaternary sea-level history and offshore wind farm development, and the overall effort certainly indicates the utility of AAR in offshore stratigraphic research.


As part of the BOEM project, we also developed an on-line database (ArcGIS Online) of AAR data in order to facilitate discussion within the working group of colleagues involved in the BOEM project.  This database is a living document that will evolve as new data and interpretations are added. 


Links to the NOAA Paleoclimate Data Site (still being populated):

General site:


AAR data files:


Search for paleodata

Enter Racemization

Browse other datasets at this link:




American Association for the Advancement of Science

American Geophysical Union          

Geochemical Society; Organic Geochemical Society

Geological Society of America         (elected GSA Fellow in 1990)

American Quaternary Association

Union of Concerned Scientists

Academy of Natural Sciences of Philadelphia



Columbia University, New York, 1966-1971; Ph. D., Geology (geochemistry).

Carnegie Inst. Washington, Geophysical Lab., pre-doctoral fellow, 1969-1971.

Swarthmore College, Swarthmore, Penna., 1962-1966; BA, Chemistry



Physical Geology; Historical Geology; Earth Resources; Quaternary Geology and Geochronology; Organic Geochemistry; General Geochemistry; Aminostratigraphy


 ~80 (~30 chaired)  1973 – present; 4 post-doc supervisions 1980- present.

Current activities:

  • Quaternary stratigraphy and geochronology of the North Carolina Coastal Plain (with USGS, North Carolina Geological Survey, and East Carolina University)
  • Age mixing of fossils in coastal sediments
  • Archiving of field guides, collection site documentation, and associated research documents at the University of Delaware Institutional Repository: http://udspace.udel.edu/handle/19716/10389
  • Online ArcGIS maps summarizing on-going data reporting and synthesis, US Atlantic coastal plain. Project supported by Bureau of Ocean Energy Management (BOEM) Atlantic Sand Assessment Program through the Delaware and South Carolina Geological Surveys.


  • Documentation and distribution of our field research collection of more than 10,000 Quaternary fossils to collaborating museums and other host institutions, including the Delaware and North Carolina Geological Surveys, the Paleontological Research Institution (Ithaca, NY), the Florida Museum of Natural History, the Academy of Natural Sciences of Drexel University, and the Natural History Museum of Los Angeles County.
  • Modeling the kinetics and diagenesis of amino acids in carbonate fossils


PUBLICATIONS 1970’s to present:

Wehmiller, J. F. and P. E. Hare (1971). Amino acid content of some samples from the JOIDES Deep Sea Drilling Program. In Initial Reports, Deep Sea Drilling Program 9, 903-905. National Science Foundation, Wash. D. C.

Wehmiller, J. F., and P. E. Hare (1971). Racemization of amino acids in marine sediments. Science 173, 907-911.

Kvenvolden, K. A., E. Peterson, J. Wehmiller, and P. E. Hare (1973). Racemization of amino acids in marine sediments determined by gas chromatography. Geochimica et Cosmochimica Acta 37, 2215-2225.

Swetland, P. J. and J. F. Wehmiller (1975).  Lipid geochemistry in the Great Marsh, Lewes, Del. In: Marine Chemistry in the Coastal Environment, T. M. Church, ed., Amer. Chem. Soc. Symposium Series 18, pp. 285-303.

Wehmiller, J. F. and M. Lethen (1975).  Saturated hydrocarbon material in Delaware Bay sediments as determined by gas chromatography.  Technical Report CMS-RANN-3-75, College of Marine Studies, Univ. Delaware (Grant Report to National Science Foundation).

Wehmiller, J. F. (1976) BOOK REVIEW: Geochemistry and the Origin of Life, edited by K. A. Kvenvolden.  Jour. Chem. Education 53, a236-a238.

Wehmiller, J. F., P. E. Hare, and G. A. Kujala (1976). Amino acids in fossil corals: racemization (epimerization) reactions and their implications for diagenetic models and geochronological  studies.   Geochimica et Cosmochimica Acta 40, 763-776.

Wehmiller, J. F., G. L. Kennedy, and K. R. Lajoie (1977). Amino acid age estimates for Pleistocene marine deposits in the Eureka-Fields Landing area, Humboldt Co., California. U. S. Geological Survey Open-file Report 77-517, pp. 1-25.

Wehmiller, J. F., K. R. Lajoie, K. A. Kvenvolden, E. Peterson, D. F. Belknap,  G. L. Kennedy, W. O. Addicott, J. G. Vedder, and R. W. Wright (1977). Correlation and chronology of Pacific coast marine terrace deposits of continental United States by fossil amino acid stereochemistry – technique evaluation, relative ages, kinetic model ages, and geologic implications. U. S. Geological Survey Open-file Report 77-680, pp. 1-196.

Wehmiller, J. F. (1977).  Amino acid studies of the Del Mar, California midden site: apparent rate constants, ground temperature models, and chronological implications. Earth and Planetary Science Letters 37, 184-196.

Wehmiller, J. F. and D. F. Belknap (1978).  Alternative kinetic models for the interpretation of amino acid enantiomeric ratios in Pleistocene mollusks: examples from California, Washington, and Florida. Quaternary Research 9, 330-348. (see also Q. R.11, 278).

Keck, R. T., R. C. Heess, J. F. Wehmiller, and D. Maurer (1978).  Sublethal effects of the water soluble fraction of Nigerian crude oil on juvenile hard clams, Mercenaria mercenaria. Environmental Pollution 15, 109-119.

Wehmiller, J. F. and W. K. Emerson (1980).  Calibration of amino acid racemization in late Pleistocene mollusks: results from Magdalena Bay, Baja California Sur, Mexico, with dating applications and paleoclimatic implications. The Nautilus 94, 31-36.

Lajoie, K. R., J. F. Wehmiller, and G. L. Kennedy (1980). Inter- and intra-generic trends in apparent racemization kinetics of amino acids in Quaternary mollusks. In: Hare, P. E.,  Hoering, T. C., and King, K., Jr.(eds.), Biogeochemistry of Amino Acids, John Wiley, New York, pp. 305-340.

Wehmiller, J. F. (1980).  Intergeneric differences in apparent racemization kinetics in mollusks and foraminifera: implications for models of diagenetic racemization. In: Hare, P. E., Hoering,  T. C., and King, K. Jr.(eds.) Biogeochemistry of Amino Acids, John Wiley, New York, pp. 341-355.

Belknap, D. F. and J. F. Wehmiller (1980).  Amino acid racemization in Quaternary mollusks: examples from Delaware, Maryland, and Virginia.  In:  Hare, P. E., Hoering, T. C., and King, K. Jr.(eds.) Biogeochemistry of Amino Acids, John Wiley, New York, pp.401-414.

Emerson, W. K., G. L. Kennedy, J. F. Wehmiller, and E. Keenan (1981).  Age relations and zoogeographic implications of late Pleistocene marine invertebrate faunas from Turtle Bay, Baja California Sur, Mexico. The Nautilus 95, 105-116.

Atwater, B. F., B. E. Ross, and J. F. Wehmiller (1981). Stratigraphy of late Quaternary estuarine deposits and amino acid stereochemistry of oyster shells beneath San Francisco Bay, California. Quaternary Research 16, 181-200.

Wehmiller, J. F. (1981).  Kinetic model options for interpretation of amino acid enantiomeric ratios in Quaternary mollusks: comments on a paper by Kvenvolden et al. (1979). Geochimica et Cosmochimica Acta 45, 261-264.

Oldale, R. N., P. C. Valentine, T. M. Cronin, E. C. Spiker, B. W. Blackwelder, D. F. Belknap, J. F. Wehmiller, and B. J. Szabo (1982).  Stratigraphy, structure, absolute age, and paleontology of the upper Pleistocene deposits at Sankaty Head, Nantucket Island, Massachusetts. Geology 10, 246-252.

Kennedy, G. L, K. R. Lajoie, and J. F. Wehmiller (1982). Aminostratigraphy and faunal correlations of late Quaternary marine terraces, Pacific coast, U. S. A. Nature 299, 545-547.

Wehmiller, J. F. and D. F. Belknap (1982).  Amino acid age estimates, Quaternary Atlantic coastal plain: comparison with U-series dates, biostratigraphy, and paleomagnetic control. Quaternary Research 18, 311-336.

McCartan, L., J. P. Owens, B. W. Blackwelder, B. J. Szabo, N. Kriausakul, R. M. Mitterer, D. F. Belknap, and J. F. Wehmiller (1982). Comparison of amino acid racemization geochronometry with lithostratigraphy, biostratigraphy, uranium-series coral dating, and magneto-stratigraphy in the Atlantic coastal plain of the southeastern United States. Quaternary Research 18, 337-359.

Wehmiller, J. F. (1982).  A review of amino acid racemization studies in Quaternary mollusks: stratigraphic and chronologic applications in coastal and interglacial sites, Pacific and Atlantic coasts, United States, United Kingdom, Baffin Island, and Tropical Islands.  Quaternary Science Reviews 1,83-120.

Keenan, E. M., L. Ortlieb, and J. F. Wehmiller (1984). Amino acid dating of Quaternary marine terraces at Bahia Asuncion, Baja California Sur, Mexico. In: Neotectonics and sea level variations in the Gulf of California area, V. Malpica-Cruz et al., eds., Univ. Nal. Auton, Mexico,  Inst. Geologia, Mexico, D. F.,  pp. 149-164.

Wehmiller, J. F. (1984).  Relative and absolute dating of Quaternary mollusks with amino acid racemization: evaluation, applications, and questions. In: Mahaney, W. C.,(ed.) Quaternary Dating Methods, Elsevier.

Wehmiller, J. F. (1984). Interlaboratory comparison of amino acid enantiomeric ratios in fossil mollusks.  Quaternary Research  vol. 22, 109-120.

Muyzer, G., P. Westbroek, J. P. M. De Vrind, J. Tanke, T, Vrijheid, E. W. De Jong, J. W. Bruning, and J. F. Wehmiller (1984).  Immunology and Organic Geochemistry.  ORGANIC GEOCHEMISTRY, vol. 6., pp. 847-865.

Amino acid racemization geochronology, In DATING YOUNG SEDIMENTS,  A. J. Hurford, E. Jager, and J.A.M. Ten Cate, Proceedings of 1985 Beijing Workshop on Dating of Young Sediments, United Nations CCOP, Bangkok, pp. 139-158, 1986.

Wehmiller, J. F. and D. F. Belknap, 1987, Aminostratigraphy of coastal U. S. Quaternary marine deposits – deciphering the timing of interglaciacial high sea levels and the thermal histories of coastal regions, IN:  CLIMATE, History, Periodicity, and Predictability, M. R. Rampino, J. E. Sanders, W. S. Newman, and L. K. Konigsson, eds., Van Nostrand Reinhold, New York,  pp. 157-165.

Wehmiller, J. F. , D. F. Belknap, and L. L. York (1987).  Comments on Corrado et al. (1986): Capabilities and limitations of applied aminostratigraphy as illustrated by analyses of Mulinia lateralis from the late Cenozoic marine beds near Charleston, S. C.: South Carolina Geology, v. 31(2), 109-117.

Wehmiller,  J. F., D. F. Belknap, B. S.  Boutin, J. E.  Mirecki, S. D.  Rahaim, and L. L.  York, 1988,    A Review of the Aminostratigraphy of Quaternary Mollusks from United States Atlantic Coastal Plain Sites.   Geol. Soc. Amer. Special Paper 227, Dating Quaternary Sediments, D. L. Easterbrook, ed. ,  pp. 69-110.

Muyzer, G., P. Westbroek, and J. F. Wehmiller (1988).  Phylogenetic implications and diagenetic stability of macromolecules from Pleistocene and recent shells of Mercenaria mercenaria.  Historical Biology.  v. 1,  pp. 135-144.

York, L. L., J. F. Wehmiller, T. M. Cronin, and T. A. Ager (1989).   Stetson Pit, Dare County, North Carolina: an integrated chronologic, faunal, and floral record of subsurface coastal  Quaternary sediments.  Palaeogeography, Palaeoclimatology, Palaeoecology.  v. 72, 115-132.  (Proceedings volume, INQUA Long Continental Records Symposium, August, 1987; G. Kukla and D. Adam, editors).

Hsu, J. T.,  E. M. Leonard, and J. F. Wehmiller (1989).  Aminostratigraphy of Peruvian and Chilean marine terraces.  Quaternary Science Reviews,  8, 255-262.

Wehmiller, J. F., J. E. Mirecki, and C. H. Fletcher (1989).  Lithostratigraphy and aminostratigraphy of the southeastern Virginia coastal plain.  IGCP 274 Guidebook, July, 1989.

Groot, J. J., K. W. Ramsey, and J. F. Wehmiller  (1990).  Ages of the Bethany, Beverdam, and Omar Formations of southern Delaware.  Delaware Geological Survey Report of Investigations No. 47, pp. 1-19.

Wehmiller, J. F. (1990)  Amino acid racemization: applications in chemical taxonomy and chronostratigraphy of Quaternary fossils, IN:  Skeletal Biomineralization: Patterns, Processes, and Evolutionary Trends, V. 1.  Joseph Carter, ed.  Van Nostrand Reinhold, New York, pp. 583-608.

Leonard, E. M. and Wehmiller, J. F. (1991).   Geochronology of marine terraces at Caleta Michilla, Northern Chile; implications for late Pleistocene and Holocene uplift.   Revista Geologica de Chile, 18(1) p. 81-86.

Wehmiller, J. F., L. L. York, D. F. Belknap, and S. W. Snyder (1992).   Theoretical correlations and lateral discontinuities in the Quaternary Amino-Stratigraphic Record of the U. S. Atlantic Coastal Plain.   Quaternary Research,  v. 38,  275-291

 Leonard, E. M. and Wehmiller, J. F. (1992).  Low uplift rates and terrace reoccupation inferred from mollusk aminostratigraphy, Coquimbo Bay area, Chile.  Quaternary Research 38, 246-259.

York, L. L., and J. F. Wehmiller (1992).  Aminostratigraphic results from Cape Lookout, N.C. and their relation to the preserved Quaternary marine record of SE North Carolina.  Sedimentary Geology  80, 279-291.

Riggs, S. R., L. L. York, J. F. Wehmiller, and S. W. Snyder (1992).  Depositional patterns resulting from high frequency Quaternary sea-level fluctuations in northeastern North Carolina.    In:  C. H. Fletcher III and J. F. Wehmiller (editors), Quaternary Coasts of the United States: Marine and Lacustrine Systems.  Society of Economic Paleontologists and Mineralogists Special Publication No.  48, pp. 141-153.

Wehmiller, J. F. (1992).  Aminostratigraphy of Southern California Quaternary marine terraces.   In:  C. H. Fletcher III and J. F. Wehmiller (editors), Quaternary Coasts of the United States: Marine and Lacustrine Systems.  Society of Economic Paleontologists and Mineralogists Special Publication No.  48, pp. 317-321.

Kennedy, G. L., Wehmiller, J. F. and Rockwell, T. K. (1992). Paleocology and paleozoogeography of late Pleistocene marine terrace faunas of southwestern Santa Barbara County, California. In:  C. H. Fletcher III and J. F. Wehmiller (editors), Quaternary Coasts of the United States: Marine and Lacustrine Systems.  Society of Economic Paleontologists and Mineralogists Special Publication No.  48, pp. 343-361.

C. H. Fletcher III and J. F. Wehmiller, editors, (1992), Quaternary Coasts of the United States: Marine and Lacustrine Systems. Society of Economic Paleontologists and Mineralogists Special Publication No. 48, pp. 1-450. 

Wehmiller, J. F. (1993).  Applications of Organic Geochemistry for Quaternary Research: aminostratigraphy and aminochronology, IN: Organic Geochemistry, M. Engel and S. Macko, eds.,  Plenum Pub. Co., pp. 755-783.

Genau, R. B., Madsen, J. A., McGeary, S., and Wehmiller, J. F. (1994).  Seismic reflection identification of Susquehanna River paleochannels on the mid-Atlantic coastal plain.  Quaternary Research, 42, 166-175.

Wehmiller, J. F., York, L. L., and Bart, M. L. (1995). Amino acid racemization geochronology of reworked Quaternary mollusks on US Atlantic coast beaches: implications for chronostratigraphy, taphonomy, and coastal sediment transport.   Marine Geology, v. 124, 303-337.

Mirecki, J. E., Wehmiller, J. F., and Skinner, A. 1995. Geochronology of Quaternary Coastal Plain deposits, southeastern Virgina, USA.   Jour. Coastal Research, v. 11(4), 1135-1144.   

Lamothe, Wehmiller, and Noller 1998; Comparison of approaches to dating Atlantic coastal plain sediments, Virginia Beach, Virginia: Citable URI: http://udspace.udel.edu/handle/19716/13228

E. M. Leonard and J. F. Wehmiller – Aminostratigraphy and chronology of marine terraces in the Coquimbo Bay area, Chile.  IGCP 367 guidebook, Nov. 1995

Martin, R.E, Wehmiller, J. F.,  Harris, M. S., and  Liddell, W. D., 1996.    Comparative taphonomy of bivalves and foraminifera from Holocene tidal flat sediments, Bahia la Choya, Sonoroa, Mexico (Northern Gulf of California); taphonomic grades and temporal resolution.   Paleobiology 22(1), pp. 80-90.

Kaufman, A., Ghaleb, B., Wehmiller,  J. F., and Hillaire-Marcel , C. (1996).  Uranium concentration and isotope ratio profiles within Mercenaria shells: geochronological implications.  Geochimica et Cosmochimica Acta 60, 3735-3746. 

Wehmiller, J. F.  (1997).  Aminostratigraphic analysis of mollusk specimens:  Cape May Coast Guard Station Borehole.  In:  Miller, K. G. and Snyder, S. W., eds.   Proceedings of Ocean Drilling Program V150X, p. 355-357.

Wehmiller, J. F. and Hall, F. R. (1997).  Amino acid racemization geochronological studies of selected Leg 155 samples.   In:  Flood, R. D., Piper, D. J. W.,  Klaus, A., and Peterson, L. C., eds.  Proceedings of Ocean Drilling Program V155, p. 375-378. 

Wehmiller, J. F.  and Miller, G. H. (1998).   Amino acid racemization, in:  Sowers, J. M., Noller, J. S. and Lettis, W. R.,  Dating and Earthquakes: Review of Quaternary geochronology and its applications to paleoseismology.  U. S. Nuclear Regulatory Commission, NUREG/CR 5562, pp. 2-307 – 2-359.

Wehmiller, J. F.  Lamothe, M., and Noller, J. S.  (1998).   Comparison of approaches to dating Atlantic Coastal Plain sediments, Virginia Beach, VA., in:  Sowers, J. M., Noller, J. S. and Lettis, W. R.,  Dating and Earthquakes: Review of Quaternary geochronology and its applications to paleoseismology.  U. S. Nuclear Regulatory Commission, NUREG/CR 5562, pp. 4-3 to 4-31.

Goodfriend, G. A., M. J. Collins, M. L. Fogel, S. A. Macko, and J. F. Wehmiller (eds.). 2000. Perspectives in Amino Acid and Protein Geochemistry. Oxford University Press, NY. xvi+366 pp.

O’Neal, M. L., Wehmiller, J. F. and Newell, W. L. (2000) Amino acid geochronology of Quaternary coastal terraces on the northern margin of Delaware Bay, southern New Jersey, USA.   Perspectives on Amino Acid and Protein Geochemistry  (G. A Goodfriend et al., editors). pp 301-319.

Wehmiller, J. F., Stecher, H. A., III,  York, L. L., and Friedman, I.  (2000)  The thermal environment of fossils: effective ground temperatures (1994-1999)  at aminostratigraphic sites, U. S. Atlantic coastal plain. Perspectives on Amino Acid and Protein Geochemistry (G. A Goodfriend et al., editors). pp 219-250.

Wehmiller, J. F. and Miller, G. H. (2000). Aminostratigraphic dating methods in Quaternary geology, in: Noller, J. S., Sowers, J. M., and Letis, W. R., eds., Quaternary Geochronology, Methods and Applications, American Geophysical Union Reference Shelf, v. 4, p. 187-222.

C. H. Fletcher, et al. (2000)  Coastal sedimentary research examines critical issues of national and global priority.  EOS, Trans AGU, 81(17), 181.

Wehmiller, J. F. and York, L. L. (2002) The Occurrence and Distribution of Fossil Shells on U. S. mid-Atlantic Beaches.  National Park Service Park Paleontology Newsletter, Vol. 6, no. 2 (summer 2002). http://www2.nature.nps.gov/grd/geology/paleo/paleo_6_2/newsletter_6_2_screen.pdf

 Wehmiller, J. F. and Farrell, K. M. .(2002).  Geochronology of Quaternary marine units, Cape Fear Arch region. Field Guide for 2002 Bald Head Island Conference.

Thomas H. O’Donnell,, Stephen A. Macko, Janel Chou, Kathy L. Davis-Hartten and John F. Wehmiller (2003). Analysis of 13C, 15N, and 34S in organic matter from the biominerals of modern and fossil Mercenaria spp.  Organic Geochemistry Volume 34, Issue 2 , February 2003, Pages 165-183

 John F. Wehmiller, Kathleen R. Simmons, Hai Cheng, R. Lawrence Edwards , Jamie Martin-McNaughton, Linda L.York, David E. Krantz , and Chuan-Chou Shen (2004) Uranium-series coral age from the US Atlantic Coastal Plain – the “80 ka problem” revisited.  Quaternary International 120, 3-14.

Muhs, D. R., Wehmiller, J. F., Simmons, K. R., and York, L. L. (2004) Quaternary sea level history of the United States, IN: A. R. Gillespie, S. C. Porter, and B. F. Atwater, eds. The Quaternary Period in the United States, pp. 147-183. Developments in Quaternary Science. Elsevier, Amsterdam.

Smerdon J. E., H. N. Pollack, V. Cermak, J. W. Enz, M. Kresl, J. Safanda, J. F. Wehmiller (2004), Air-ground temperature coupling and subsurface propagation of annual temperature signals, J. Geophys. Res., 109, D21107, doi:10.1029/2004JD005056. 

Mallinson, D., Riggs, S., Thieler, E. R., Culver, S., Farrell, K., Foster, D., Corbett, R., Horton, B.and Wehmiller, J. (2005) Late Neogene and Quaternary evolution of the northern Albemarle Embayment (mid-Atlantic continental margin, USA) Marine Geology 217 97-117

Barbour-Wood, S., Krause, R., Kowalewski, M., Wehmiller, J., and Simoes, M. (2006). Aspartic acid racemization dating of Holocene brachiopods and bivalves from the southern Brazilian shelf, South Atlantic. Quaternary Research 66, 323-331.

Smerdon J. E., H. N. Pollack, V. Cermak, J. W. Enz, M. Kresl, J. Safanda, J. F. Wehmiller (2006), Daily, seasonal, and annual relationships between air and subsurface temperatures.  J. Geophys. Res., 111, DO7101, doi:10.1029/2004JD005578, 2006.

Gardner, N., Hall, B., and Wehmiller, J.  (2006) Pre-Holocene Raised Beaches at Cape Ross, Southern Victoria Land, Antarctica,  Marine Geololgy 229, 273-284.

O’Donnell, T. H., Macko, S. A., and Wehmiller, J. F. (2007) Stable carbon isotope composition of amino acids in modern and fossil Mercenaria. Organic Geochemistry 38, 485-498..

Parham, P., S. R. Riggs, S, J. Culver, D. J. Mallinson, and J. F. Wehmiller (2007) Quaternary depositional patters and sea level fluctuations, northeastern North Carolina. Quaternary Research 67, 83-99.

Stephen J. Culver, Kathleen M. Farrell, David J. Mallinson, Benjamin P. Horton, Debra A. Willard, E. Robert Thieler, Stanley R. Riggs, Scott W. Snyder, John F. Wehmiller, Christopher E. Bernhardt, Caroline Hillier (2008). Micropaleontologic record of late Pliocene and Quaternary paleoenvironments in thenorthern Albemarle Embayment, North Carolina, U.S.A.Palaeogeography, Palaeoclimatology, Palaeoecology 264 (2008) 54–77

David J. Mallinson, Stephen J. Culver, Stanley R. Riggs, E. Robert Thieler, David Foster,John Wehmiller, Kathleen M. Farrell, Jessica Pierson (2010). Regional seismic stratigraphy and controls on the Quaternary evolution of the Cape Hatteras region of the Atlantic passive margin, USA. Marine Geology 268, 16–33.

Wehmiller, J.F., et al.,  (2010) Aminostratigraphy of surface and subsurface Quaternary sediments, North Carolina coastal plain, USA, Quaternary Geochronology  5, 459-492. doi:10.1016/j.quageo.2009.10.005.

Krause et al. (2010).  Quantitative comparisons and models of time-averaging in bivalve and brachiopod shell accumulations. Paleobiology 36(3), 428-452.

Culver, S. J., Farrell, K. M., Mallinson, D. J., Willard, D. A., Horton, B. P., Riggs, S. R., Thieler, E. R., Wehmiller, J. F., Parham, P., Snyder, S. W., Hillier, C. (2011).  Micropaleontologic record of Quaterenary paleoenvironments in the Central Albemarle Embayment, North Carolina, U. S. A.  Palaeogeography, Palaeoclimatology, Palaeoecology 305, 227-249.

Wehmiller et al., 2012. Calibration of amino acid racemization (AAR) kinetics in United States mid-Atlantic Quaternary mollusks using 87Sr/86Sr analyses: evaluation of kinetic models and estimation of regional Late Pleistocene temperature history. Quaternary Geochronology 7, 21-36.

Wehmiller, J. F. , 2013. United States Quaternary coastal sequences and molluscan racemization geochronology – what have they meant for each other for the past 45 years?   Quaternary Geochronology 16, 3-20.

Simonson, A. E., Lockwood, R., and Wehmiller, J. F., 2013.  Three approaches to radiocarbon calibration of amino acid racemization in Mulinia lateralis from the Holocene of Chesapeake Bay, USA  Quaternary Geochronology, 16: 62-72.

Wehmiller, J., 2013.  Interlaboratory comparison of amino acid enantiomeric ratios in Pleistocene fossils.  Quaternary Geochronology, 16: 173-182.

Miller, K. et al. 2013.  Pleistocene Sequence Stratigraphy of the shallow continental shelf, offshore New Jersey: Constraints by IODP Leg 313 coreholes.   Geosphere 9, 74-95.

Farrell, K. M. et al. 2013. Graphic logging for interpreting process-generated stratigraphic sequences and aquifer/reservoir potential: with analog shelf to shoreface examples from the Atlantic coastal plain province, U. S. A.  Journal of Sedimentary Research, 83, 723-745.

Tuysuz, O., et al. 2013.  Geology and morphology of the Antakya Graben between the Amik Triple Junction and the Cyprus Arc.  Geophysical Research Abstracts, vol 15, EGU2013-2064-2. 

Wehmiller and Pellerito, 2015.  An evolving database for Quaternary aminostratigraphy.  GeoResJ, 6, 115-123.  Available at this link: http://www.sciencedirect.com/science/article/pii/S2214242815000170

Wehmiller and Pellerito, 2015.  Open-File Report at the Delaware Geologic Survey containing the UD amino acid database and associated files.   http://www.dgs.udel.edu/publications/ofr50-database-quaternary-coastal-geochronologic-information-atlantic-and-pacific-coast

Wehmiller et al., 2015 Geological Society of America Annual Meeting Poster: Racemization-inferred age distribution of mollusks in the US Atlantic margin coastal system. Available on the GSA meeting web site: https://gsa.confex.com/gsa/2015AM/webprogram/Paper263915.html

Culver, S. J. et al., 2016.  Micropaleontologic record of Pliocene and Quaternary paleoenvironments in the southern Albemarle Embayment, North Carolina, U.S.A.  Palaeogeography, Palaeoclimatology, Palaeoecology 457, 360-379.

Wehmiller, J.F., Ramsey, K. E., and Thieler, E. R. 2017.  Paired racemization and radiocarbon analysis of Pleistocene and Holocene shells, US Atlantic coastal plain: implications for shell geochronology, Pleistocene sea level history, and paleoclimate.  Geological Society of America, Southeastern Section, Richmond VA.  March 2017


Wehmiller, J. F. 2017.  On-line databases for Atlantic coastal plain aminostratigraphic data.  Geological Society of America, Southeastern Section, Richmond VA.  March 2017



Wehmiller et al., 2019  SE GSA  Oral presentation:  Beach shells: https://gsa.confex.com/gsa/2019SE/webprogram/Paper326646.html

Wehmiller et al., 2019 SE GSA  Poster: https://gsa.confex.com/gsa/2019SE/webprogram/Paper326300.html

Long, J.H.; Hanebuth, T.J.J.; Alexander, C.R., and Wehmiller, J.F., 2021. Depositional environments and stratigraphy of Quaternary paleochannel systems offshore of the Georgia Bight, southeastern U.S.A. Journal of Coastal Research, 37(5), 883–905. Coconut Creek (Florida), ISSN 0749-0208.  https://doi.org/10.2112/JCOASTRES-D-20-00088.1

Wehmiller et al., 2021. Molluscan aminostratigraphy of the US Mid-Atlantic Quaternary coastal system: Implications for onshore-offshore correlation, paleochannel and barrier island evolution, and local late Quaternary sea-level history.  Quaternary Geochronology 66, 101177

2022  Wehmiller and Miller  Long Term preservation of amino acid racemization laboratory comparison samples.  Quaternary Geochronlogy  vol 70, 2022  101290